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Similarly, a clade of fungi interacting with primitive fungus‐gardening ants—thought to be ancestral to the more derived leaf‐cutter symbionts—appears instead to be a more recent acquisition from free‐living stock.

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Reciprocal specialization and vertical symbiont cotransmission are thought to promote a pattern of largely synchronous coevolutionary diversification in attines.

Here we test this hypothesis by inferring the first time‐calibrated multigene phylogeny of the lepiotaceous attine cultivars and comparing it with the recently published fossil‐anchored phylogeny of the attine ants.

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